L-R: Clarkia bottae (Farewell-to-spring), Blennosperma nanum (Blennosperma), Fritillaria biflora (Chocolate lily), Toxicoscordion fremontii (Star lily), Calochortus kennedyi (Desert mariposa lily)

In the following names, the stressed vowel is the one preceding the stress mark. It is not always easy to ascertain where such stress should be placed, especially in the case of epithets derived from personal names. I have tried to follow the principle of maintaining the stress of the original name as outlined in the Jepson Manual, and have abandoned it only when it was just too awkward. In the case of some names, I have listed them twice, reflecting either some disagreement or conflict in the rules of pronunciation, some uncertainty on my part as to the correct pronunciation, or that simply sometimes there is no single correct pronunciation. In other instances, the way I record it is just that which sounds right to my ear. Where no credit is given for photos, they are in public domain mostly from Wikipedia.
  • tabernaemonta'ni: after Jacob Theodor von Bergzabern (Jacobus Theodorus Tabernaemontanus) (1520-1590). The
      following is from Stearn's Dictionary of Plant Names: "...personal physician to the Count of the Palatine at Heidelberg, West Germany. He was born in the town of Bergzabern in the Palatinate region of Germany and was a student of the German botanist Hieronymus Bock.  He Latinized his name as Tabernaemontanus [which means Tavern in the mountains] and is also commemorated by species named for him in Amsonia, Potentilla and Scirpus. He was the author of a celebrated herbal Neuw Kreuterbuch (1588-1591), of which the illustrations were issued separately at
    Frankfurt-am-Main in 1590 under the title Eicones Plantarum. The woodcuts were mostly copied from those in other herbals but make an attractive book. The London printer, John Norton, acquired them from the Frankfurt printer Nicholaus Bassaeus and used them in 1597 to illustrate Gerard's Herball." Many of the websites that mention von Bergzabern have to do with beer and brewing, a subject he was apparently very interested in. He studied at the University of Heidelberg, was married three times and had eighteen children. He spent a lifetime botanizing and practicing medicine and has been called the ‘Father of German botany.’ His work provided material for the better-known Herball of John Gerard (1597).  He supported himself by serving as a court physician to a number of German nobles such as Philip III, Count of Nassau-Weilburg and Marquard von Hattstein, bishop of Speyer. The genus Tabernaemontana was created in his honor by Charles Plumier and the name was adopted and published by Carl Linnaeus in 1753. (ref. Scirpus tabernaemontani)
  • Taenia'therum: from the Greek tainia, "a band or ribbon," and ather, "an awn," referring to the flat or ribboned awn of the lemma. The genus Taeniatherum was published by Sergei Arsenjevic Nevski in 1934. (ref. genus Taeniatherum)
  • Tage'tes: named after the Etruscan god Tages who supposedly emerged from the earth as it was being ploughed and was imbued with the power of divination. The genus Tagetes was published by Carl Linnaeus in 1753. (ref. genus Tagetes)
  • tageti'na: possibly from genus Tagetes. (ref. Navarretia tagetina)
  • tahoen'sis: of or from the area of Lake Tahoe. (ref. Carex tahoensis)
  • Tam'arix: the Latin name for this plant derived from the Tamaris River in Spain. The genus Tamarix was published by Carl Linnaeus in 1753. (ref. genus Tamarix)
  • tanacetifo'lia: tansy-leaved, with leaves like Tanacetum. (ref. Machaeranthera tanacetifolia, Phacelia tanacetifolia)
  • Tanace'tum: from the medieval Latin name tanazita, in turn derived from Greek athanasia, "immortality," of uncertain application to this taxa. The genus Tanacetum was published by Carl Linnaeus in 1753. (ref. genus Tanacetum)
  • tandilen'sis: named for the Sierra de Tandil, near the center of Buenos Aires Province, Argentina, where this taxon is native. (ref. Agrostis tandilensis)
  • taraxacifo'lia: with leaves like genus Taraxacum.
  • taraxaco'ides: having a resemblance to Taraxacum. (ref. Leontodon taraxacoides)
  • Tarax'acum: medieval name traceable through Arabic to the Persian talkh chakok, meaning "bitter herb. The Desktop Guide to Herbal Medicine by Brigitte Mars (2009) states “Opinions differ on the origin of dandelion’s genus name, Taraxacum. Some believe that it derives from the Persian talkh chakok, “bitter herb.” Others propose that it derives from the Greek taraxos, “disorder,” and akos, “remedy.” Still others believe it could be derived from the Greek taraxia, “eye disorder,” and akeomai, “to cure,” as the plant was traditionally used as a remedy for eyes.” The online Free Dictionary says “from Medieval Latin, from Arabic tarakhshaqūn, "wild chicory," perhaps of Persian origin. And the 1894 Vol. 4 of the Agricultural Gazette of New South Wales says “Though it is a plant which must have been well known to the ancients, no distinct reference to it can be traced, either In the classics of Greece or Italy, although a plant mentioned by Theophrastus is thought to be it. The word taraxacum is usually considered to be of Oriental origin, probably meaning “wild lettuce”, and we first meet with it in the works of Arabian physicians, who regarded it as a sort of wild endive. It is thus mentioned by Rhazes in the Tenth and by Avicenna in the Eleventh Century. Some commentators consider it to be one of the bitter herbs eaten with the Passover lamb by the Israelites when leaving Egypt.”  The dandelion has formerly gone by the Latin names of Taraxacum densleonis and Leontodon taraxacum which is what Linnaeus called it. Both names have reference to the French dent de lion and the Latin dens leonis, meaning "tooth of the lion," and are an allusion apparently to the jagged edges of the plant’s leaves. The dandelion had been known and utilized for centuries, and it had a widespread use in 18th century herbal pharmacology. Dandelion concoctions were common in European drugs of the early 19th century. Linnaeus called the common dandelion Leontodon Taraxacum, the specific name being adopted from an old time classical name. Leon is Latin for lion and -odon is Greek for tooth, so he thus used two linguistic sources for his genus name, and took the name Taraxacum from a Middle Eastern language, perhaps with the thought of illustrating the plant’s wide usage and appeal. The genus Taraxacum was published by Friedrich Heinrich Wiggers in 1780. (ref. genus Taraxacum)
  • Tarax'ia: so named because it had leaves similar to Leontodon taraxacoides. The Greek word taraxia means "eye disorder." Other words which this epithet may be related to are taraxis, "disorder" or "tumult," and a-taraxis, "tranquility," or "absence of disorder." The name was originally published by John Torrey and Asa Gray, and clearly has some relationship to the name of the dandelion, Taraxacum. (See Taraxacum). I thank a correspondent, Eric Neville, for inquiring about the name Taraxia and for sending me a source of information about it, which led me to investigate further this name and that of the related name Taraxacum. The genus Taraxia was published by Thomas Nuttall in 1893. (ref. genus Taraxia)
  • tardiflor'a: late-flowering or developing.
  • tarminia'na: apparently named after a Columbian agrnomist and professor named Tarmín de Jesus Campos Espinosa (1947- ) who was working with fruit researchers in Columbia and introduced them to the study of passion fruits. (ref. Passiflora tarminiana)
  • tatar'ica: from the common name of this species, Russian blue lettuce, I infer that that the name tatarica refers to the Tatars, a primarily Turkic peoples found in areas of the former U.S.S.R. Lonicera tatarica is native to Siberia and the assignment of the name of the native Lactuca tatarica was based on a plant from Siberia described by J. G. Gmelin as 'Sonchus foliis lanceolatis sessilibus dentatis, floribus corymbosis, caulibus glabris'. Linnaeus renamed it Sonchus tataricus and C.A. Meyer transfered it to Lactuca in 1831. (ref. Lactuca tatarica, Lonicera tatarica)
  • taur'icum: of or from the Crimean region of the Mediterranean, which in ancient times was referred to as Taurica Chersonesus. (ref. Onopordum tauricum)
  • Tau'schia: named after Ignaz Friedrich Tausch (1793-1848), a Bohemian professor of botany, naturalist, plant collector, director of the garden of the Duke of Canal de Malabaillas in Prague, and author of Hortus Canlius, seu Plantarum Rariorum…Icones et Descriptiones in 1823. Tausch was born in Bohemia to a master brewer, Josef Tauch. In his youth, Tausch visited a Piaristengymnasium, a higher educational institution, in the town of Schlackenwerth, sparking his curiosity in botany. From 1809 to 1812, he attended the Charles University in Prague, where he studied philosophy, medicine, and natural sciences. In 1815 he was appointed as a professor of economic and technical botany and from then until 1826 he served as a professor of botany at the botanical garden of Emmanuel Canal at Prague. He was a member of the Academy in Turin, and in 1843 became president of the Bohemian Horticultural Society. He discovered at least eleven species of plants, including Rhizobotrya alpina and Saxifraga hostii. He was the taxonomic authority of many botanical species. Plants bearing the specific epithet of tauschii are named in his honor, and in 1835, Diederich Franz Leonhard von Schlechtendal named the genus Tauschia in the Apiaceae after him. He died at Prague. The genus Tauschia was published by Diederich Franz Leonhard von Schlechtendal in 1834. (ref. genus Tauschia)
  • taxifo'lia: with leaves like those of the genus Taxus or yew. (ref. Suaeda taxifolia)
  • Ta'xus: the Latin name for the yew tree. The genus Taxus was published by Carl Linnaeus in 1753. (ref. genus Taxus)
  • tay'lorii: after Dean William Taylor (1948- ), a California botanist who is particularly interested in mountain regions like the Sierras and the White Mountains. He studied with G. Ledyard Stebbins and received his doctoral degree from UC Davis. He has been a lecturer at San Francisco State University, University of Colorado at Boulder, University of California at Santa Cruz, and a research botanist at Oregon State University. He was the first Vice-President of the California Botanical Society and is currently on the Rare Plant Scientific Advisory Committee of the California Native Plant Society. He has been deeply involved in the conservation of endangered plants. (ref. Erythronium taylorii)
  • tazet'ta: apparently an Italian vernacular name from tazza, "a small cup," referring to the form of the corona, which is a petal-like or crown-like structure between the petals and stamens in some flowers. (ref. Narcissus tazetta)
  • Teco'ma: abbreviated from the Mexican name tecomaxochitl. The genus Tecoma was published by Antoine Laurent de Jussieu in 1789. (ref. genus Tecoma)
  • Tecomar'ia: takes its name from Tecoma, a species which it closely resembles. The genus Tecomaria was published by Édouard Spach in 1840. (ref. genus Tecomaria)
  • tecopen'sis: named after Tecopa, a hot springs community southeast of Death Valley National Park. (ref. Cordylanthus tecopensis)
  • tector'um: relating to the roofs of houses, this was a name used by Linnaeus for various Swedish plants that grew on thatched roofs. (ref. Bromus tectorum, Crepis tectorum)
  • Teesdal'ia: after Robert Teesdale (1740-1804), an English botanist, seedsman and horticulturist from Yorkshire. He was a friend of James Edward Smith, and contributed to Smith and Sowerby's English Botany, a major publication of British plants comprising a 36 volume set, issued in 267 monthly parts over 23 years from 1791 to 1814. He also put out Catalog of the More Rare Plants in the Neighborhood of Castle Howard. He was a founder member (1788) of the Linnean Society and gardener to the Earl of Carlisle at Yorkshire. The genus Teesdalia was published in 1812 by William Townsend Aiton. The genus Teesdalia was published by William Townsend Aiton in 1812. (ref. genus Teesdalia)
  • tegetario'ides: like tegetarius. This taxon is similar to Astragalus kentrophyta var. tegetarius. (ref. Astragalus tegetarioides)
  • tegetar'ius: from the Latin tegetarius, "a mat maker," from teges or tegetis, "a covering, mat." (ref. Astragalus kentrophyta var. tegetarius)
  • tehamen'se: of or from Tehama County in northern California north of Sacramento. (ref. Hesperolinon tehamense)
  • Telli'ma: anagram of Mitella. The genus Tellima was published by Robert Brown in 1823. (ref. genus Tellima)
  • telmatei'a: of or referring to wet meadows or pools. (ref. Equisetum telmateia ssp. braunii)
  • tembloren'se: same as next entry. (ref. Eriogonum temblorense)
  • temblorien'sis: named for the Temblor Range in south-central California bordering on the San Joaquin Valley and the Carrizo Plain. (ref. Clarkia tembloriensis)
  • temulen'ta/temulen'tum: from the Latin temulus for "drunken, nodding, top-heavy." (ref. Festuca temulenta)
  • ten'ax: gripping, tenacious or sticky in one sense, and firm, persistant or stubborn in another, from teneo or tenere, "to hold." (ref. Iris tenax, Xerophyllum tenax)
  • tenebro'sa: growing in shady places.
  • tenel'la/tenel'lum/tenel'lus: from the Latin meaning "quite delicate, dainty." (ref. Chorispora tenella, Gentianella tenella, Hesperocnide tenella, Osmadenia tenella, Physaria tenella, Lithophragma tenellum, Plagiobothrys tenellus, Psilocarphus tenellus)
  • te'ner: slender, tender, soft. (ref. Erigeron tener, Astragalus tener)
  • tene'rrima/tener'rimus: very slender. (ref. Gilia tenerrima, Poa tenerrima, Rorippa tenerimma, Sonchus tenerrimus)
  • ten'ue: see tenuis. (ref. Galium porrigens var. tenue)
  • tenui-: prefix indicating the characteristic of being slender.
  • tenuicau'lis: with fine stems.
  • tenuiflor'a/tenuiflor'us: with fine or delicate flowers. (ref. Gilia tenuiflora, Carduus tenuiflorus)
  • tenuifo'lia/tenuifo'lius: with finely-divided, slender leaves. (ref. Calystegia macrostegia ssp. tenuifolia, Diplotaxis tenuifolia, Ipomopsis tenuifolia, Malacothrix saxatilis var. tenuifolia, Potamogeton alpinus ssp. tenuifolius)
  • tenuilo'ba: with finely-divided or slender lobes. (ref. Mirabilis tenuiloba)
  • tenuiloba'tum: see tenuiloba above. (ref. Solanum tenuilobatum)
  • tenu'ipes: with a slender stalk. (ref. Hydrophyllum tenuipes)
  • tenu'is: slender. (ref. Castilleja tenuis, Lessingia tenuis, Nemacladus tenuis, Malperia tenuis, Orcuttia tenuis)
  • tenuisec'ta/tenuisec'tum: thinly or narrowly cut. (ref. Verbena tenuisecta, Cirsium arizonicum var. tenuisectum)
  • tenuis'sima/tenuis'simus: very slender. (ref. Iris tenuissima, Potamogeton pusillus var. tenuissimus)
  • tenuitu'ba: from the Latin tuba, "a trumpet" and the prefix tenui- which means "slender or narrow" referring to the flowers. (ref. Ipomopsis tenuituba)
  • tephro'des: from the Greek tephros for "ash-colored" in reference to the leaves. (ref. Helianthus niveus ssp. tephrodes)
  • terebinthifo'lius: this is a bit of a puzzle. Terebinthinaceus means pertaining in some way to turpentine, and turpentine (at least the Chian variety) derives from the turpentine tree, Pistacia terebinthus, a member of the sumac family and a native of the Mediterranean region, specifically but perhaps not limited to the island of Chios. So perhaps terebinthifolius means "having leaves like the turpentine tree." (ref. Schinus terebinthifolius)
  • terebinth'inus: resembling or appearing like turpentine. (ref. Cymopteris terebinthinus)
  • te'res: cylindrical, circular in cross-section. (ref. Diodia teres)
  • tereticor'nis: with terete or cylindrical horns. (ref. Eucalyptus tereticornis)
  • teretifo'lia/teretifo'lium/teretifo'lius: terete-leaved, that is, with leaves that are smooth and cylindrical, usually circular in cross-section. (ref. Ericameria teretifolia, Erysimum teretifolium, Chrysothamnus teretifolius)
  • terna'ta/terna'tum: with parts in groups of three, referring often to the leaflets. (ref. Keckiella ternata, Eriogonum ternatum)
  • terna'tea: of the island of Ternate in the Moluccas. (ref. Clitoria ternatea)
  • tern'ipes: from the Latin terni, "three," and the suffix -pes, referring to the stalk, or foot of something, hence, "three-stalked." (ref. Aristida ternipes)
  • terraci'na: uncertain meaning, but possibly having something to do with terraces (?), or a reference to Terracina, a seaside resort area between Rome and Naples, which perhaps makes more sense since this species is originally from the Mediterranean. (ref. Euphorbia terracina)
  • terres'tris: in Latin means "on land." (ref. Brodiaea terrestris ssp. kernensis, Tribulus terrestris)
  • tescam'nis: I was contacted by Diana Hurlbut at the New York State Museum where the author of this taxon was a curator to the effect that the protologue for this species in Rhodora states that "the specific epithet is derived from tesca, 'desert,' [or wilderness, wastelands] and amnis, 'swift-flowing river,' in allusion to the species’ typical habitat along arid-region streams." According to Bill Jennings, one of the authors, "Most specimens are from lower elevations near rivers, in the Intermountain Region." (ref. Platanthera tescamnis)
  • teso'ta: corruption of the Spanish tieso meaning "stiff or firm". Tesota is a southwest Native American name for this tree. (ref. Olneya tesota)
  • tessella'ta: tessellate or checkered, patterned like a mosaic, in allusion to the arrangement of warts on the back of the nutlets. (ref. Amsinckia tessellata)
  • testicula'tus: like testicles. (ref. Ranunculus testiculatus)
  • Tetracoc'cus: "four seeds," from the Greek tetra, "four," and kokkos, "a kernel, grain, or berry," (from Edmund Jaeger's A Source-Book of Biological names and Terms) because of the four-lobed or four-seeded ovary in the original species, T. dioicus.  My original information followed Philip Munz, who used the word kakkos for "fruit," but I have been kindly informed by Dr. Stavros Kakkos of my mistake, and in checking other sources find the proper word source to have been kokkos.  Thank you, Dr. Kakkos. The genus Tetracoccus was published by Georg Engelmann in 1885. (ref. genus Tetracoccus)
  • Tetrady'mia: from the Greek tetra, "four," and dymos, "together," from the four-flowered heads of the first known species of this genus. The genus Tetradymia was published by Augustin Pyramus de Candolle in 1837. (ref. genus Tetradymia)
  • tetrago'na: probably the same as the next entry. (ref. Crassula tetragona)
  • Tetrago'nia: four-angled. The genus Tetragonia was published by Carl Linnaeus in 1753. (ref. genus Tetragonia)
  • tetragonio'ides: like genus Tetragonia. (ref. Tetragonia tetragonioides, or literally the Tetragonia that looks like Tetragonia)
  • tet'rahit: the Missouri Plants website suggests that this means "four-parted." The name Tetrahit was apparently used as a generic name by French physician and botanist Matthias de l'Obel (1538-1616), but I have no further information on its derivation or meaning. (ref. Galeopsis tetrahit)
  • tetramer'a/tetramer'es: from tetras, "four," and meris, "part," thus meaning "with four parts. (ref. Phacelia tetramera, Tetradymia tetrameres)
  • tetrancis'tra: from the Greek tetra, "four," and ankistron, "fish-hook," thus meaning 4 fish-hooks for the 3-4 hooked central spines. (ref. Mammillaria tetrancistra)
  • tetran'dra: from the Greek tetra, tetras, "four" and aner, andros, "man, male, stamen." (ref. Primula tetrandra)
  • Tetraneur'is: from the Greek tetras, "four," and neuron, "nerve." The genus Tetraneuris was published by Edward Greene in 1898. (ref. genus Tetraneuris)
  • tetraphyl'lum: four-leaved (ref. Polycarpon tetraphyllum)
  • Tetrap'teron: Greek for "four-winged." The genus Tetrapteron was published by Warren Lambert Wagner and Peter C. Hoch in 2007. (ref. genus Tetrapteron)
  • tetrasper'ma: four-seeded. (ref. Vicia tetrasperma)
  • Teu'crium: there seems to be a lot of confusion about this name, although most of the confusion may be on my part alone. The Jepson Manual simply says an ancient Greek name. The Botanary website says "Named for Teucer, a Trojan king who used the plant as a medicine; or possibly for Dr. Teucer, a botanist and physician." Stearns' Dictionary of Plant Names says: "Possibly named for Teucer, first king of Troy." The Columbia Encyclopedia gives two alternate meanings: "1. Ancestor and king of the Trojans, who are also called the Teucri. He was the father-in-law of Dardanus. 2. Son of Telamon and Hesione. He was the greatest archer in the Trojan War and a faithful comrade of his half brother, the Telamonian Ajax. When he returned home he was banished by his father, who mistakenly thought that Teucer was responsible for the death of Ajax. Teucer went to Cyprus, where he founded the town of Salamis and ruled as king." Umberto Quattrocchi states: "From the Greek teukrion, possibly for Teucer (Teukros), the founder of the town of Salamis in Cyprus." Encyclopedia Mythica says: "Teucer was the son of the river Scamander and the nymph Idaea, and was the legendary ancestor of the Trojans; hence the Trojans are often called 'Teucrians.' He should not be confused with the Teucer who was the son of Telamon and the brother of Ajax, and who fought against Troy during the Trojan War." And Wikipedia states that Teucer was "The son of Hesione and Telamon, Teucer fought with his half-brother, Ajax the Great, in the Trojan War and is the legendary founder of the city Salamis on Cyprus." It appears that there were two separate figures, Teucrus, who was the ancestor of the Trojans, and the Greek Teucer who fought in the Trojan War and founded Salamis. However, another website stated that the Teucer who was the founder ancestor of the Trojans also fought in the Trojan War against the Greeks, and I don't see how that would have been possible. As to which of these figures is honored by the name Teucrium, Pliny wrote that Teucer discovered Teucrium during the same period in which Achilles discovered Achillea, so he would be refering to the one who fought in the Trojan War. The genus Teucrium was published by Carl Linnaeus in 1753. (ref. genus Teucrium)
  • texa'na/texa'num/texa'nus: of or from, or somehow relating to, Texas. (ref. Bergia texana, Erodium texanum, Nuttallanthus texanus)
  • texen'sis: same as previous entry. (ref. Carex retroflexa var. texensis)
  • tex'tilis: from the Latin textilis, "woven or braided." (ref. Juncus textilis)
  • thacheria'num: after Olive Day Thacher (1891-1971), member of the California Botanical Society, recipient in 1915 of a BS in botany from the University of California, and daughter of Sherman Day Thacher who in 1889 founded the Thacher School in Ojai, which was then known as the School for Out-of-Door Life and Study. She collected Ribes thacherianum on Santa Cruz Island in 1921. (ref. Ribes thacherianum)
  • thalia'na: after Johann Thal (1542-1583), German physician and botanist. He was born in Erfurt, Thuringia in central Germany, the son of a Protestant pastor. He was first schooled in Erfurt then went to the monastery school at Ilfeld (1558-1561), a then famous school under the direction of Michael Neander, a man Thal worshipped as a second father after the early death of his own father. Soon after entering that school, he had become enamored of botany and the natural world and had determined 72 grass species in the surrounding area and created his own herbarium. He then studied medicine at the University of Jena beginning in 1561 and practiced his profession in Stendal and then in Stolberg as a doctor in the city (1572), Whether he actually received any kind of medical degree is unclear. He worked for five years producing a compilation of the plants of the Harz Mountains and northern Thuringia region. While authors who preceded Thal had limited themselves to medicinally active plants, Thal's work was unique because it included all occurring plants. He first discovered what came to be called Arabidopsis thaliana in the Harz Mountains. He originally called it Pilosella siliquosa, and it went through several name changes before finally being named in his honor in 1842. This species has turned out to be an incredibly significant and important model organism in developmental biology. In 1583 he had a carriage accident and fractured his lower leg, dying as a result of his severe injuries three weeks later. His work Sylvia Hercynia was published posthumously in 1588. (ref. Arabidopsis thaliana)
  • Thalic'trum: from thaliktron, a name used to describe a plant with divided leaves, and a name given to the genus by Dioscorides, the Greek physician and pharmacologist who wrote the Materia Medica, which remained the leading pharmacological text for sixteen centuries. The genus Thalictrum was published by Carl Linnaes in 1753. (ref. genus Thalictrum)
  • Thamnos'ma: from the Greek for "odorous shrub." The genus Thamnosma was published by John Torrey and John Charles Frémont in 1845. (ref. genus Thamnosma)
  • -thamnus: a shrub. (ref. genera Chrysothamnus, Malacothamnus)
  • thap'sus: David Hollombe provides this derivation: "Nikander, Theocritus and Theophrastus wrote about Thapsos, a plant from Thapsos, Sicily. The root was used to produce a yellow dye. Some sources claim it was Cotinus coggygria." The name Verbascum thapsus was published by Linnaeus in 1753. (ref. Verbascum thapsus)
  • -theca: cover, case, container. (ref. genus Heterotheca, also Spergularia macrotheca)
  • Thelesper'ma: from the Greek thele, "nipple," and sperma, "seed," and referring to the roughened achenes. The genus Thelesperma was published by Christian Friedrich Lessing in 1831. (ref. genus Thelesperma)
  • Thelypo'dium: from the Greek thelys, "female," and pus, "foot," from the presence of the stipe or stalk above the receptacle. The genus Thelypodium was published by Stephan Friedrich Ladislaus Endlicher in 1839. (ref. genus Thelypodium)
  • Thelyp'teris: from the Greek thelys, "female," and pteris, "fern." The genus Thelypteris was published by Casimir Christoph Schmidel in 1763. (ref. genus Thelypteris)
  • theophras'ti: named after the Greek philospher and botanist Theophrastos (371-c.287 B.C.). The following is quoted
      from a website called Theophrastus of Eresos: "Around 320 BC the Greek philosopher Theophrastus begins the science of botany with his books De causis plantarum/The Causes of Plants and De historia plantarum/The History of Plants. In them he classifies 500 plants, develops a scientific terminology for describing biological structures, distinguishes between the internal organs and external tissues of plants, and gives the first clear account of plant sexual reproduction, including how to pollinate the date palm by hand. Theophrastus (or Theophrast or Theophrastos)
    (371 or 372 -287/286) BC, the son of Melantas, born in Eresos on Lesbos, was a student of Aristotle and succeeded him as a director of the Lyceum in Athens. He took over the philosophy of Aristotle in parts reshaping, commenting, and developing it in an original way. His thinking leads to empirism by means of observation, collection, and classification. He was around 35 years the director of the Lyceum and he was a teacher of up to 2000 students. His true name was Tyrtamos of Eresos. Due to his oratory talents he was nicknamed Euphrastos, the well-spoken, eventually to become famous as Theophrastus, divine spoken. Having joined Plato’s Academy at the age of 17 he soon fell to Aristotle’s spell and accompanied him, still a young man, in his self-chosen exile on the Troad then on his home island Lesbos. He then disappeared from the record for three or more years – during which time some believe he traveled far, to Crete and Libya to come again at Aristotle’s side in Stageira. From there on he never again left his master except for his short last exile, succeeding him as the headmaster of the peripatetic school until his death in 287, at the venerable age of 85. He is said to have been a congenial chap, sworn bachelor and gourmet, and to have died of the sequels of the wedding party of one of his pupils. The main innovation of Theophrastus is his attempt to find a connection between the 'first principles' (the intelligible world, ratio) and the perceivable objects of nature; this distinction remains the main motive of occidental philosophy during the next two millenia with different solutions. Theophrastus is also called 'father of botany' and can be regarded as the founder of ecology, too. He described the origin of plants from seeds, he carried out germination experiments, discussed the influence of abiotic habitat factors on plants, the ecology of domestic plants, pollination of plants with the example of the fig, he invented a growth form terminology which is still valid (root, shoot). He described more than 500 species and varieties of plants from lands bordering the Atlantic and Mediterranean. He classified plants into trees, shrubs, under shrubs, and herbs. He noted that some flowers bear petals whereas others do not, and observed the different relative positions of the petals and ovary. In his work on propagation and germination, Theophrastus described the various ways in which specific plants and trees can grow: from seeds, from roots, from pieces torn off, from a branch or twig, or from a small piece of cleft wood." (ref. Abutilon theophrasti)
  • Theres'ia: named for his wife Therese Auguste Weichardt Koch (1818-1886) by German physician and botanist Karl Heinrich Emil Koch in 1849. The genus Theresia was published by Karl Heinrich Emil Koch in 1849. (ref. genus Theresia)
  • therma'le/therma'lis: of warm springs. (ref. Gnaphalium canescens ssp. thermale, Phacelia thermalis)
  • thermop'ola: David Hollombe at first thought this was for Thermopolis, Wyoming, but then found out that "the type locality was on the Snake River near the southern entrance to Yellowstone and the range was originally thought of as a narrow band from Yellowstone to Salt Lake City, all far to the west of Thermopolis, but I think there is still a reference to hot springs, just different ones." (ref. Artemisia arbuscula ssp. thermopola)
  • Thermop'sis: from the Greek thermos for lupine and opsis, "like," bearing yellow lupine-like flower heads. The genus Thermopsis was published by Robert Brown in 1811. (ref. genus Thermopsis)
  • Thing'ia: named for Thing of the Addams Family. Mark Hershkovitz in the journal Phytoneuron published this generic name in 2019. He says: “The generic name metaphorically refers to one of the colloquial names for the type species, ‘dead man’s fingers.’ Thus, the genus is named for the literary figure, Thing, of the Addams Family cartoon, television series, and movie. ‘Thing’ is an animated but (at least in the movie) disembodied human hand. This suggests that the hand derives from a deceased human, hence possesses “dead man ́s fingers.” The generic combination also suggests an appropriate vernacular name, ‘ambiguous thing,’ reflecting the ambiguity of the status of this species following cladistic classification of Montiaceae.” The genus Thingia was published by Mark A. Hershkovitz in 2019. (ref. genus Thingia)
  • Thlas'pi: from the Greek thlaein, "to crush," from the flattened silicle, and the Greek name for a cress. The genus Thlaspi was published by Carl Linnaeus in 1753. (ref. genus Thlaspi)
  • thom'asii: after George Henry Thomas (1816-1870), United States Army officer stationed at Fort Yuma about
      1850, and later as a general a Civil War hero called the Rock of Chickamauga. He was born of French Huguenot ancestry at Newsom’s Depot, Virginia on an upper class plantation which eventually reached a size of almost 700 acres and included 24 slaves. After his father died, he and his family had to flee their home and hide in the woods during the slave uprising of Nat Turner in 1931. This episode appeared to create in him a negative view of slavery, although he continued to own slaves. Thomas was appointed to the United States Military Academy at West Point, New
    York, in 1836 by Congressman John Y. Mason, where one of his roommates was William Tecumsah Sherman. He served in the Seminole Wars in Florida and then during Mexican-American War. In 1851 he was appointed as a cavalry and artillery instructor at West Point, where he established a close professional and personal relationship with another Virginia officer, Lt. Col. Robert E. Lee. One of his students who received his recommendation for assignment to the cavalry was Jeb Stuart. Despite his southern heritage chose to side with the Union during the Civil War, in which he had a distinguished career. He had an uncomfortable relationship with General Grant which may partially explain why he did not seem to get the credit for his war service that he deserved, but he was widely regarded as the next best general to Grant and Sherman on the Union side. He had an interest in botany and plant collecting, and it was because of plants he collected near Fort Yuma that John Torrey honored him with the specific epithet thomasii. He died of a stroke in 1870. (ref. Eriogonum thomasii)
  • thompson'iae: after Ellen Louella Powell Thompson (1843-1911), called "Nellie," American naturalist and botanist, and an active advocate for women's suffrage. She was born in New York and was a founding member of the Women's Anthropological Society of America, Washington DC. She was the sister of John Wesley Powell and having a degree in botany she accompanied her husband, Almon Harris Thompson, on an expedition through the Escalante Wilderness in 1872. Thompson had been appointed by Powell to lead the expedition after he left to return to Salt Lake City to visit his wife and new child. They charted the course of the Escalante River which was the last named river in the United States. At one point the expedition climbed a pass between two peaks: Mt. Ellen and Mt. Pennell. "Prof" Thompson named Mt. Ellen after his wife, "Nellie." She was the first person to do botanical studies in the region. On his expedition of the year before, Powell had named one of his boats the Nellie Powell. Another brother was William P Powell, superintendent of Washington DC public schools. She attended Wheaton College in Wheaton, Illinois, in the mid-1850s and taught school from the age of 16. In 1862 she was married to professor, topographer, explorer, geologist, geographer and Civil War veteran Almon Harris Thompson, who was a colleague and friend of her brother John. Ellen Thompson accompanied her husband on expeditions to map the western United States. During this period, she made friends with members of a number of Indian tribes, learning the language of the "Pah Utes" and studying their customs. When her husband entered the Army, she took her husband's position as superintendent of schools, and spent one summer caring for the sick and wounded soldiers. She had at least five taxa named after her. (ref. Penstemon thompsoniae, Peteria thompsoniae)
  • thorn'ei: after Robert Folger Thorne (1920-2015), "Bob" to his friends, Professor of Botany Emeritus at Claremont
      Graduate University and Rancho Santa Ana Botanical Garden, botanist and authority on angiosperm evolution, and former President of Southern California Botanists. The following is quoted from the April/May newsletter of the Orange County Chapter CNPS: "He is retired from his position as curator and taxonomist at the Rancho Santa Ana Botanic Garden, although he continues to conduct research. He is a member of the San Diego chapter of CNPS. He continues also as an active member of Southern California Botanists, where he served previously as
    member of the board, vice-president, and president. As a young man - he is now 84 - he served as a B-24 bomber navigator in World War II. During a mission over Austria, his plane was badly shot up. Despite the fact that the plane was riddled with flak, causing a fire in one engine, the crew managed to parachute over the only partisan-held island (Vis) in the Dalmatian chain, now part of Croatia. (Had he landed on any other island, he and his crewmates would have been prisoners of war). But Dr. Thorne’s military career did not end at that point - after the crew was returned to Italy he flew 29 more missions. Dr. Thorne became hooked on botany when he took required science courses at Dartmouth College. Abandoning plans to become a linguist, he went on to earn an M.S. in 1942 and a Ph.D in 1949 from Cornell University. After graduation, he spent thirteen years at the University of Iowa, as an assistant, associate, then full professor. He became friends with Peter Raven (another giant in the field) during a five-month stint as a researcher at the Royal Botanic Gardens at Kew and the British Museum of Natural History in London. Dr. Raven was then the curator at Rancho Santa Ana, and the Garden accepted his recommendation to appoint Dr. Thorne as his successor when the former moved on to Stanford. During his tenure there (from 1962 to 1987), Dr. Thorne was also Professor of Botany at Claremont Graduate School, now Claremont Graduate University. At that time, famed cactologist Lyman Benson was curator of the herbarium at Pomona College. When Dr. Benson retired, the Pomona College collection was donated to Rancho Santa Ana, and Dr. Thorne became curator of the combined collection. Before and during those years, Dr. Thorne traveled extensively while collecting specimens for herbaria. He spent 1½ years in Australia as a Fulbright researcher studying primitive angiosperms in the rain forests of the region, including New Caledonia, New Guinea, and Tasmania. Based on his travels in Mexico, he has prepared a flora of the high country of the Sierra San Pedro Martir and is working on a flora of the California floristic region of northwestern Baja California. His California publications include the Flora of the Santa Ana Mountains and the Flora of the Santa Rosa Plateau (both with Earl Lathrop), the Flora of the Higher Ranges of the Eastern Mojave (with Jim Henrickson and Barry Prigge) and the Flora of Santa Catalina Island. His numerous other works include two chapters written for the voluminous Terrestrial Vegetation of California, treatments of aquatic monocots in the Jepson Manual and many papers on biogeography, vernal pools, other plant communities of California as well as floras of areas in New York, Iowa, Virginia, Georgia, and Florida. Among other awards and honors and memberships Dr. Thorne has received is the Asa Gray Award from the American Society of Plant Taxonomists. He received two Fulbright scholarships and a National Science Foundation Senior Postdoctoral Fellowship while at the University of Iowa. He has been a Fellow of the Linnaean Society of London and was honored with foreign membership in the Royal Society of Denmark." (ref. Eriogonum thornei, Delphinium variegatum ssp. thornei)
  • Thorneochlo'a: see previous entry. The genus Thorneochloa was published in 2019 by Konstantin Romaschenko, Paul M. Peterson, and Robert John Soreng. The genus Thorneochloa was published by Konstantin Romaschenko, Paul M. Peterson and Robert John Soreng in 2019. (Ref. genus Thorneochloa)
  • Thu'ja: from the Greek name thuia or thyia, for a kind of juniper or other resinous tree. The genus Thuja was published by Carl Linnaeus in 1753. (ref. genus Thuja)
  • Thunber'gia: after Carl Peter Thunberg (1743-1828), the ‘Father of South African botany.’ Stearn’s Dictionary
      says: He was a "...Swedish botanist, a student of Linnaeus who was pursuaded by Dutch lovers of new plants to enter the service of the Dutch East India Company as a doctor and send back plants from Japan to Europe. He travelled in South Africa and Japan and became professor of botany at Uppsala." He was born in Jönköping, Sweden, and attended Uppsala University where he was one of the students of Carl Linnaeus and graduated in 1767. Linnaeus encouraged him in 1770 to travel to Paris, Amsterdam and Leiden to deepen his knowledge of botany, medicine and natural history.
    Linnaeus’ Swedish pupil Johannes Burman convinced Thunberg to travel to the Indies to collect plants for the Leiden botanical garden, which he did for the Dutch East India Company in 1771. He was also experiencing a powerful attraction to South Africa, which he reached in 1772. He was there for three years and regularly undertook field trips and journeys into the interior of South Africa, during which time he not only collected specimens of flora and fauna but also studied the culture of the native people of Western South Africa who were called the Khoikhoi. At some point he met Francis Masson, a Scots gardener who had come to Cape Town to collect plants for the Royal Gardens at Kew. While in South Africa he graduated in absentia from the University of Uppsala as Doctor of Medicine. In 1775 he left the Cape for Batavia and travelled on to Japan.  At first restricted to a small artificial island in the Bay of Nagasaki by order of the Shogun, he was gradually allowed limited access to the city, and began trading medical knowledge for botanical information and the ability to collect specimens. He taught local doctors about treating syphilis and in turn learned about acupuncture. Wikipedia goes on to say: “In both countries, Thunberg's knowledge exchange hence led to a cultural opening-up effect which too manifested itself also in the spread of universities and boarding schools which taught knowledge on the other culture. For this reason, Thunberg has been given the title of being "the most important eye witness of Tokugawa Japan in the eighteenth century." “Due to his scientific reputation, Thunberg was given the opportunity in 1776 to accompany the Dutch ambassador M. Feith to the shogun's court in Edo, today's Tokyo. During that journey, the Swede was given the chance to collect a great number of specimen of plants and animals and likewise to talk to Japanese locals in the villages they traversed on their way. It is in this time that Thunberg wrote two of his scientific masterpieces, the Flora Japonica (1784) and the Fauna Japonica (1833). The latter was completed by the German traveller Philipp Franz von Siebold who visited Japan between 1823 and 1829. Yet, von Siebold based the Fauna Japonica on Thunberg's notes which he carried with him all the time in Japan.” In November 1776 he left Japan and travelled first to Java, then to Ceylon, before returning to Europe via South Africa in 1778. He made a short trip to London to meet Sir Joseph Banks, and then upon arriving back in Sweden, was appointed in 1781 professor of medicine and natural philosophy at the University of Uppsala. He published his Flora japonica in 1784, and in 1788 he began to publish his travels. He completed his Prodromus plantarum in 1800, his Icones plantarum japonicarum in 1805, and his Flora capensis in 1813. He had been elected a member of the Royal Swedish Academy of Sciences and then in 1823 associated member of the Royal Institute of the Netherlands. He died near Uppsala in 1818. The genus Thunbergia was published in 1780 by Anders Jahan Retzius. The genus Thunbergia was published by Anders Jahan Retzius in 1776. (ref. genus Thunbergia)
  • thur'beri: named after Dr. George Thurber (1821-1890), called the most accomplished horticulturist in America, and botanist and quartermaster of the Mexican Boundary Survey, 1850-1854. Dr. Thurber was professor of botany and horticulture at Michigan Agricultural College 1859-1863 and editor of the American Agriculturalist from 1863 until his death in 1890. He was born in Providence, Rhode Island and studied at the Union Classical and Engineering School, but did not graduate. He apprenticed with a local pharmacist and then set up in business with a partner, while at the same time giving lectures in chemistry at the Franklin Society of Providence. He was largely self-taught as a botanist, and like so many others was stimulated in his botanical explorations by his desire to learn about medicinal plants. In 1850 he was offered the position of botanist on the U.S. Boundary Commission which had been tasked to survey the boundary between the United States and Mexico, and he remained with the Commission until 1854. The collections he made, including the curious Pilostyles, form the subject of Asa Gray's Plantae Novae Thurberianae. Returning to academia, he earned a graduate degree in chemistry from Brown University and was appointed, probably through the influence of Dr John Torrey, to the United States Assay Office in New York. Wikipedia says: “He left the Assay Office in 1856 and held a succession of teaching positions at Cooper Union, the College of Pharmacy of the City of New York, and Michigan Agricultural College, before assuming the position of editor of American Agriculturist, which he occupied for 22 years. As editor, he not only wrote regular columns on botany and horticulture but was also dedicated to exposing business and professional frauds, and the many swindlers and charlatans circulating within the agricultural community. After his death, Garden and Forest magazine praised him for having elevated the standing of the American agricultural and horticulture press more than any other writer of his time. His larger publications included American Weeds and Useful Plants, the graminology of the botany of California, and the botany entries for Appleton's New American Encyclopaedia. Thurber, who suffered most of his life from a debilitating rheumatism contracted in Mexico, did not live to complete his monograph on American grasses on which he laboured for many years. He became a corresponding member of the Royal Horticultural Society of London in 1886, six years after his tour of the European continent, during which he met many leading botanists and horticulturalists. In America, he was a member of the New Jersey Horticultural Society, the Philadelphia Academy of Sciences, the New York Academy of Sciences, and the American Pomological Society, and received an honorary doctorate from the University Medical College of New York. After the death of his good friend Dr Torrey in 1873, Thurber served as president of the Torrey Botanical Club for seven years. Two different genera of plants were given the same name in dedication to him, though neither is in current usage. Thurberia A. Gray (1855) in the Malvaceae is a synonym of Gossypium. The later homonym Thurberia Bentham (1881) in the Poaceae is an illegitimate name for Limnodea L.H. Dewey.” (ref. Centrostegia thurberi, Eriogonum thurberi, Lepidium thurberi, Penstemon thurberi, Petalonyx thurberi, Pilostyles thurberi)
  • thurberia'na/thurberia'num: see previous entry. (ref. Agrostis thurberiana, Achnatherum thurberianum)
  • Thymophyl'la: from the Greek thymos, "thyme," and phyllon, "leaf," meaning "thyme-leaved." The genus Thymophylla was published by Mariano Lagasca y Segura in 1816. (ref. genus Thymophylla)
  • thysanocar'phus: like platycarphus, there are two possibilities here: (1) from Greek thusanos, "fringe," and karphos, "a dry splinter, twig, straw," in turn from karpho, "to dry up or wither," referring to the pappus or to the scales of the involucre; (2) it is also possible that this is just an alternate spelling of 'thysanocarpus' meaning "fringed fruit." (ref. Orochaenactis thysanocarphus)
  • thyrsiflor'a: with flowers in a thyrse, which according to Harris and Harris's Plant Identification Terminology is "a compact cylindrical or ovate panicle with an indeterminate main axis and cymose sub-axes." (ref. Lysimachia thyrsiflora)
  • Thysanocar'pus: from the Greek words thusanos, "fringe," and karpos, "fruit," hence "fringed fruit." The genus Thysanocarpus was published by William Jackson Hooker in 1830. (ref. genus Thysanocarpus)
  • Tiarel'la: from the Greek tiara, tiaras for "a small tiara", a Persian head-dress worn on great occasions. The genus Tiarella was published by Carl Linnaeus in 1753. (ref. genus Tiarella)
  • tiburonen'sis: I think this refers to the town of Tiburon in Marin County. (ref. Calochortus tiburonensis)
  • Tidestro'mia/tidestro'mii: named for Swedish-born American botanist Ivar Theodor Tidestrøm (1864-1956), professor of botany at Catholic University, author of Flora of Utah and Nevada (1925) and Flora of Arizona and New Mexico (1941) with Sister Teresita Kittell. He apparently also did work in Maryland and Virginia. The genus Tidestromia was published by Paul Carpenter Standley in 1916. (ref. genus Tidestromia, also Astragalus tidestromii, Lupinus tidestromii)
  • tiehm'ii: after amateur plant collector and professional bellhop and limo driver Arnold Gerard Tiehm (1951- ) of Reno, Nevada, author of Nevada Vascular Plant Types and Their Collectors, published by the New York Botanical Garden in 1996. Tiehm has had the rare good luck to have discovered nineteen new plants, six of which bear his name, all found on days off in his native Nevada "backyard." He is also the author of a biography of Per Axel Rydberg, and according to David Hollombe, is at work on a biography of Amos Arthur Heller. (ref. Arabis tiehmii, Juncus tiehmii)
  • tilin'gii: named after Heinrich Sylvester Theodor Tiling (1818-1871), a German-Russian physician and botanist
      for a Russian-American company in Sitka who also collected plants in California and Nevada for various European botanical gardens. He was born in Latvia and studied medicine in Dorpat from 1838 to 1844. He became a physician at the Russian North American Co. in Ayan, Siberia from 1845 through 1851. He described all of the plants in the area and published an account of it with the director of the botanical gardens in St. Petersburg, Russia, Eduard August von Regel (Florula Ajanensis). In 1852 he and his family travelled from Ayan to Sakhalin, Kamchatka, Sitka,
    Hawaii, Tahiti, around Cape Horn through the Atlantic Ocean to Kronstadt and finally back to Riga. There he practiced for a couple of years before moving to Wenden, Lativia, where he practiced from 1854 to 1863. He resituated in 1863 to Sitka, Alaska where he was a surgeon until 1868 before moving on to San Francisco and then later to Nevada City, California. Alaska had been purchased by the U.S. government in 1867. At some point he became an American citizen. The collection and categorization of plants, and the recording of meteorological data were clearly the two major preoccupations of his life, and he was responsible for the introduction into Europe of a number of previously unknown species. More than 20 species were named for him. He wrote about his "trip around the world" in a book whose title translates as A journey around the world from West to East through Siberia and the Pacific and Atlantic seas. He died in Nevada City, California, in 1871, and his wife died in Riga in 1876. (Photo credit: Geni) (ref. Erythranthe tilingii) (Photo credit: Geni)
  • tillae'a: for the Italian botanist Michael Angelo Tilli (1655-1740), physician, professor of botany and Fellow
      of the Royal Society of London, author of Catalogus plantarum horti Pisani (1723), and from 1685 until his death Praefectus (the person in charge) of the Botanical Garden of the University of Pisa. He was born in Castelfiorentino, Italy, and became a doctor of medicine in 1677 at the University of Pisa. Wikipedia says: “He opened a practice in Florence and worked from 1681 as a doctor on the ship routes leading from Florence to Mallorca and Menorca . In 1683 he traveled to Constantinople and Adrianople to treat the son-in-law of Sultan Mehmed IV. During the return
    journey that took him through the Aegean Sea , he made notes on the natural history of the Aegean Sea. In 1685 he became a professor of botany at the University of Pisa and director of the botanical garden founded there by Luca Ghini. To expand the plant collection, he received many rare species from the Botanical Garden in Amsterdam." The genus Tillaea was named by Pier Antonio Micheli. (ref. Crassula tillaea)
  • timor'um: a modern Latin dictionary defines timoris as "fear, alarm, dread; a terror" but I don't know whether this has anything to do with the meaning here. David Hollombe says that this name is from the Latin root timoros or timoreo meaning "to help, lend aid, give succour" but again the application is unclear. (ref. Eriogonum grande var. timorum)
  • tinctor'ia/tinctor'ius: used in dyeing, and usually used to refer to a plant that when broken exudes some kind of stain. (ref. Collinsia tinctoria, Collomia tinctoria, Carthamus tinctorius)
  • tinc'tus: colored. (ref. Astragalus purshii var. tinctus)
  • tingita'nus: of Tangiers, whose ancient name was Tingis. (ref. Lathyrus tingitanus)
  • tioga'na: the only geographical feature I know of with the name of Tioga is Tioga Pass on the road going into Yosemite National Park from the eastern side, and that is what I think this name refers to since this taxon's range is the High Sierras and the White & Inyo Mts. (ref. Carex tiogana, Gentiana newberryi var. tiogana)
  • Tiqui'lia: a native South American name for a flower of this genus. I thank San Diego County botanist Michael Simpson for sending me the following from Richardson., T. 1977, Monograph of the genus Tiquilia (Coldenia, sensu lato), Boraginaceae: Ehretioideae, Rhodora 79: 467-572.: "Ruiz and Pavon (1799) described Lithospermum dichotomum,
    which they collected in Peru, and subsequently, Persoon (1805) established the genus Tiquilia to accommodate this taxon, the generic name being derived from the vernacular name, reported
    by Ruiz and Pavon (1799) as "Tiquil-tiquil". It is possible that the cited vernacular name is simply a corruption of the Quechua word for flower, "t'ika" (Pers. comm., Helen Barler. April, 1975; Lira, 1973)." The genus Tiquilia was published by Christiaan Hendrik Persoon in 1805. (ref. genus Tiquilia)
  • ti'ti: after Frank H. Titus (1850-1908). "Graduated from Medical College of the Pacific, San Francisco, CA, 1876; resident physician of St. Luke's Hospital S.F., 1890-92; Supt. S.F. City & Co. Hospital, 1894-95; proprietor of a S.F. pharmacy, 1896; U.S. Army surgeon in the Philippines 1898-1902 (shortened from Cantelow & Cantelow)." (ref. Astragalus tener var. titi)
  • tobi'ra: the Dave's Garden Botanary site says that this is from the Japanese name for this plant, the common name being Japanese pittosporum. (ref. Pittosporum tobira)
  • Tofield'ia: after Thomas Tofield (1730-1779), a British botanist and civil engineer, bom at Wilsic Hall near
      Wadworth (4 miles south of Doncaster) on 18 December, 1730, the only surviving son of Thomas Tofield (1695-1747) and his wife Elizabeth Atkinson. He attended William Burrow's School at Chesterfield and in 1747 went up to Trinity College, Cambridge, taking his B.A. in 1751. Returning to Wilsic, he began an intensive study of the local flora. This activity brought him into contact with William Hudson who in the late 1750s was working on a new British flora based on the Linnean system. Tofield sent him many specimens. In Hudson's classic Flora Anglica (1762),
    Tofield, along with John Blackstone, was singled out for special acknowledgement. He continued to send botanical data, much of which was included in the enlarged second edition of the Flora (1778). It appears that botany was only his avocation, his chief interest and occupation being as a hydrological engineer and waterways surveyor, involved mainly with the drainage of various low grounds and the diversion of rivers. The genus Tofieldia was published by William Hudson in 1778. (ref. genus Tofieldia) (Photo credit: Geni)
  • Tol'miea/tol'miei: named after William Fraser Tolmie (1812-1886), a Scottish physician with the Hudson Bay
      Company first at Fort Vancouver then elsewhere. He was born in Inverness, Scotland, and his mother died when he was only three. He was educated at Inverness Academy and Perth Grammar School. He had been interested in botany as a young child, and as so many others had found a connection between botany and medicine. This brought him in contact with the famed botanist, William Jackson Hooker, then professor of botany at the University of Glasgow, and with Dr John Scouler, who had made a voyage to Fort Vancouver (Vancouver, Wash.) in a Hudson’s Bay
    Company supply ship in 1825.  Thanks to an uncle’s encouragement he took to medical studies and was at the University of Glasgow from 1829 to 1831 although he did not receive a degree and was not an MD, but instead was a licentiate of the Faculty of Physicians and Surgeons of Glasgow, a body independent of the university. In 1832, when the Hudson's Bay Company was looking for medical officers for the Columbia District of the Pacific Northwest, Tolmie signed a five-year contract to serve in the dual capacity of clerk and surgeon. He continued working for the Hudson Bay Company in one capacity or another until he retired in 1871. He was the first white man to reach the summit of Mount Rainier, now known as Tolmie Peak. He was manager of the Puget Sound Agricultural Company, the HBC's farming subsidiary. He was elected to the House of Assembly of Vancouver Island as member for Victoria in January 1860. He was re-elected in 1863, and remained a member until Vancouver Island was annexed by the mainland colony of British Columbia in 1866. He became involved in education on Vancouver island serving for several years first on the General Board of Education and then as Chairman of the Board. He was also a member of the Legislative Assembly of British Columbia, (1874-1878). One of his sons, Simon Fraser Tolmie, became the Prime Minister of British Columbia. The genus Tolmiea was published by John Torrey and Asa Gray in 1840. (ref. genus Tolmiea, also Allium tolmiei, Saxifraga tolmiei) (Photo credit: Find-a-Grave)
  • Tol'pis: Umberto Quattrocchi says, "Origin unknown, possibly from the Greek tolype, 'a ball of wool, lump,' referring to the fruiting capitula [i.e. the small flowering heads]; some suggest from Crepis." The genus Tolpis was published by Michel Adanson in 1763. (ref. genus Tolpis)
  • tomentel'la/tomentel'lus: with small woolly hairs, diminutive of tomentosa, the oak species that bears this name being somewhat tomentose. (ref. Frangula californica ssp. tomentella, Quercus tomentella)
  • tomento'sa/tomento'sum/tomento'sus: densely covered with matted wool or short hair, tomentose. (ref. Amsonia tomentosa, Angelica tomentosa, Arctostaphylos tomentosa, Cassia tomentosa, Lessingia glandulifera var. tomentosa, Opuntia tomentosa, Pickeringia montana var. tomentosa, Phoradendron tomentosum, Ceanothus tomentosus)
  • tomentulo'sa: slightly tomentose.
  • tomp'kinsii: after chemical engineer and photographer Philip Weber Tompkins (1873-1972). The following is quoted from the California Academy of Sciences: "Philip Tompkins was born in San Anselmo, California. He graduated from the University of California in 1894. An analytical chemist and chemical engineer, he was a founder of the San Francisco chemical firm of Curtis and Tompkins where he continued to work until two years before his death (on 6 December 1972 in San Anselmo, California.) An avid photographer, he explored Utah, Colorado, and Arizona. Tompkins aided in discovering and recording the "Lost Valley of the Goblins" in Utah (1949). His article, "Goblin Valley, Recent History and Need for Protection" accompanied by many of his photographs of the area appeared in National Parks Magazine (October-December 1954). As an expression of appreciation to the Botany Department of the California Academy of Sciences, and a memorial to Alice Eastwood, he funded the Tompkins, Tehipite Botanical Expedition of the Sierra Nevada, California. An account of this journey was published in Leaflets of Western Botany by John Thomas Howell (1958). Tompkins also assisted in the publication of A Flora of Lassen Volcanic National Park, California (1961). Tompkins was a California Academy of Sciences member (1930) and Academy lecturer (1953 "Sections of South-Central Utah", 1955 "Southern Utah Scenes"). His extensive collection of slides, photographs, and negatives were donated to the Academy Library (1957, 1963)." His photographs also appeared in Mesa Land: The History and Romance of the American Southwest by Anna Wilmarth Ickes. (ref. Carex tompkinsii)
  • Tonel'la: derivation unknown. The genus Tonella was published by Thomas Nuttall in 1868. (ref. genus Tonella)
  • Tones'tus: anagram of Stenotus. The genus Tonestus was published by Aven Nelson in 1904. (ref. genus Tonestus)
  • ton'sum: smooth.
  • Toril'is: name used by Adanson in 1763, meaning obscure. Umberto Quattrocchi says "A meaningless name or possibly from toreo "to bore through, to pierce," referring to the prickled fruit. The genus Torilis was published by Michel Adanson in 1763. (ref. genus Torilis)
  • torqua'ta: presumably from the Latin torquatus, "adorned with a necklace." (ref. Eucalyptus torquata)
  • Tor'reya/torreya'na: named after John Torrey (1796-1873), a professor of chemistry and one of the giants of North
      American botany who described hundreds of plants brought or sent back by such explorers as John C. Fremont, William Emory, Charles Wilkes, Joseph Nicollet, Howard Stansbury and Charles Pickering, and sent back east also by the Mexican Boundary Expedition and the Pacific railroad survey expeditions, and who named many California species.  Born in New York, he began observing and collecting plants while still a youth. His father was appointed Fiscal Agent for the State Prison of New York when he was 15, and it was there that he met Amos Eaton, a pioneer in
    the field of natural science education. Eaton encouraged the boy's natural inclinations in the sciences, and at the age of 21 Torrey was selected to prepare A Catalog of the Plants Growing Within Thirty Miles of New York. A year later, he received a medical degree and opened a practice, although he continued to spend a great deal of his leisure time on botany. He became Professor of Chemistry, Mineralogy and Geology, and also Assistant-Surgeon, at West Point in 1824, and then Chair of Chemistry and Botany at the College of Physicians and Surgeons in New York three years later. At one time he was teaching at Columbia, Princeton and West Point simultaneously. His great contribution was not as a collector, but rather as a taxonomist, and he was the first American to move away from the Linnean system of plant classification.  Using John Lindley's system of arranging plants by families, he became of the first botanists to apply this to a major work, A Compendium of the flora of the northern and middle states. His interest in plants expanded to the Great Plains and the western Rocky Mountains after receiving the collections of Dr. Edwin James, botanist of the Long expedition of 1820. He was appointed State Botanist of New York in 1836 and in 1843 published A Flora of the State of New York. He became the mentor and lifelong friend and colleague of Asa Gray, who came to New York as a young student in the mid-1830s to study under the eminent Torrey and became his chemistry lab assistant. The two men co-authored Flora of North America, a major botanical work that was based at least in part on the descriptions and specimens of new plants sent east by Thomas Nuttall, but which was never completely finished during their lifetimes. His name was given to the Torrey pine in California by plant explorer Charles Parry who first found it.  A year before his death, he visited Parry in the Rocky Mountains of Colorado, and the latter was able to show him not only the peak he had named in Torrey's honor but also many living plants that he had described from dead specimens fifty years before. On that same trip he stayed with his friend George Engelmann in St. Louis and met John Muir in California. He was President of the American Association for the Advancement of Science, and twice President of the New York Lyceum of Natural History. As though all of his other occupations were not enough, he also served as supervisor of the New York Mint, and despite being one of the greatest American botanists, I have read that he never was able to make a living as a botanist, and he always considered that chemistry was his true profession. He founded the Botanical Club of New York which later became the Torrey Botanical Club, and his own collection eventually went to the New York Botanical Garden. The genus Torreya was published by George Arnott Walker Arnott in 1838. (ref. genus Torreya, also species Melica torreyana, Pinus torreyana, Suaeda torreyana)
  • tor'reyi: see torreyana above. (ref. Amaranthus torreyi, Atriplex torreyi, Collinsia torreyi var. wrightii, Epilobium torreyi, Juncus torreyi, Lathyrus torreyi, Lomatium torreyi, Lycium torreyi, Malacothrix torreyi, Mentzelia torreyi, Mimulus torreyi, Phyllospadix torreyi, Plagiobothrys torreyi)
  • Torreyochlo'a: after botanist John Torrey (1796-1873), see torreyana above. The genus Torreyochloa was published by George Lyle Church in 1949. (ref. genus Torreyochloa)
  • torticul'mis: with twisted stems. (ref. Eleocharis torticulmis)
  • tortifo'lia: from the Latin for "twisted leaf." (ref. Xylorhiza [formerly Machaeranthera] tortifolia)
  • tortil'is: twisted. (ref. Chylismia walkeri ssp. tortilis)
  • tortuo'sum/tortuosus: winding, very twisted. (ref. Jasminum tortuosum, Streptanthus tortuosus)
  • tournefor'tii: after Joseph Pitton de Tournefort (1656-1708), author of Institutiones Rei Herbariae published
      in 1700 and a celebrated French botanist who was the first to properly define genera and was an important forerunner of Linnaeus. He was born in Aix-en-Provence and studied at the Jesuit convent there. He was another of those who studied both botany and medicine. Wikipedia says: “After two years collecting, he studied medicine at Montpellier, but was appointed professor of botany at the Jardin des Plantes in Paris in 1683. During this time he travelled through Western Europe, particularly the Pyrenees, where he made extensive collections. Between 1700 and
    1702 he travelled through the islands of Greece and visited Constantinople, the borders of the Black Sea, Armenia, and Georgia, collecting plants and undertaking other types of observations. He was accompanied by the German botanist Andreas Gundelsheimer (1668–1715) and the artist Claude Aubriet (1651–1742). His description of this journey was published posthumously (Relation d'un voyage du Levant), he himself having been killed by a carriage in Paris; the road on which he died now bears his name (Rue de Tournefort).” Another botanist who had been one of de Tournefort’s students and accompanied him on his voyages was Charles Plumier. De Tournefort’s major work was his Eléments de botanique, ou Méthode pour reconnaître les Plantes published in 1694. Again from Wikipedia: “The classification followed was completely artificial, and neglected some important divisions established by earlier botanists, such as John Ray's separation of the phanerogams from the cryptogams, and his division of the flowering plants into monocots and dicots. Overall it was a step backwards in systematics, yet the text was so clearly written and well structured, and contained so much valuable information on individual species, that it became popular amongst botanists, and nearly all classifications published for the next fifty years were based upon it. Tournefort is often credited with being the first to make a clear distinction between genus and species. Though he did indeed cluster the 7,000 plant species that he described into around 700 genera, this was not particularly original. Concepts of genus and species had been framed as early as the 16th century, and Kaspar Bauhin in particular consistently distinguished genera and species. Augustus Quirinus Rivinus had even advocated the use of binary nomenclature shortly before Tournefort's work was published.” His herbarium collection of 6,963 specimens was housed in the Jardin du Roi in Paris, now part of the Muséum national d'histoire naturelle. He died the year after Linnaeus was born. (ref. Brassica tournefortii)
  • Townsend'ia: named for David Townsend (1787-1858), an amateur botanist. He grew up on a farm in northern Chester County, Pennyslvania, and moved to West Chester in 1810 where he worked as a clerk in the office of the Register of Wills. He was Chester county commissioner, the head bank cashier of the National Bank of Chester County. In 1826 he was the founding member of the Chester County Cabinet of Natural Science, which became West Chester University. He was a devoted and respected botanist who collected over 700 plants in the area near where he lived. Townsend was a life-long friend and business associate of Dr. William Darlington (see Darlingtonia). From the West Chester Patch August 2018: “In a relativity short period of time, Townsend established himself as respected botanist. He planted a large 'herbarium' on his property, and with his 'habits of close observation,' as his obituary described it, he was soon sharing his insights of native plants with eminent botanists in England. His assistance in identifying native asters, for instance, led the director of Kew Gardens [William Jackson Hooker] to name an entire genus in Townsend's name. Perhaps most important to Townsend was the success of his book, Florula Cestrica, which documented all the flora he had discovered in Chester County.” The genus Townsendia was published by William Jackson Hooker in 1834. (ref. genus Townsendia)
  • Toxicoden'dron: means "poison tree." The genus Toxicodendron was published by Philip Miller in 1754. (ref. genus Toxicodendron)
  • Toxicoscor'dion: Scott Earle of Larkspur Books provides the following: "Scordion is a Greek word for garlic, thus "poisonous garlic" from the rather remote resemblance of the death camases to Allium sativum," and I thank him for his contribution. The genus Toxicoscordion was published by Per Axel Rydberg in 1903. (ref. genus Toxicoscordion)
  • trachycar'pus: from the Greek trachys, "rough," and karpos, "fruit." (ref. Plagiobothrys trachycarpus)
  • trachycau'lus: rough-stemmed. (ref. Elymus trachycaulus)
  • trachygo'num: rough-kneed or rough-joined. (ref. Eriogonum trachygonum)
  • trachyphyl'la: rough-leaved. (ref. Festuca trachyphylla)
  • trachysper'ma: from the Greek trachys, "rough," and sperma, "seed." (ref. Polanisia dodecandra ssp. trachysperma)
  • tra'cyi/Tracyin'a: after Joseph Prince Tracy (1879-1953), title examiner of Eureka and botanist who amassed a collection of some 30,000 specimens from north-western California and adjacent Oregon which subsequently became part of the University Herbarium of the University of California. He studied at the University of California at Berkeley with such luminaries as Harvey M. Hall, J. Burtt Davy, Willis Jepson, and William Setchell, and graduated in 1903. Financial constraints prevented him from continuing his formal studies, but he continued collecting in Humboldt County and surrounding areas and became a leading authority on the local flora. He was a member of the California Botanical Society, the Save-the-Redwoods League, the American Association for the Advancement of Science, the American Geographic Society, and the California Academy of Sciences. (ref. Collomia tracyi, Eriastrum tracyi, Lomatium tracyi, Lupinus tracyi, Penstemon tracyi, Romanzoffia tracyi, Sanicula tracyi, also genus Tracyina)
  • Tradescan'tia: after John Tradescant (1608-1662) (called John Tradescant the Younger), English gardener to
      King Charles I. He was born in Meopham, Kent, England, the son of John Tradescant the Elder. He entered The King's School, Canterbury at the age of 11. In 1634, after a period of apprenticeship, he was admitted a freeman of the Worshipful Company of Gardeners. In 1637 he sailed to Virginia to collect seeds and plants for English gardens. Among the seeds he brought back were great American trees like the Magnolia, Bald Cypress and Tulip tree, the yucca plant, and garden plants such as phlox and asters. Some sources say that Tradescant visited Virginia on three
    occasions but I have not been able to verify that with dates. In 1638, he was appointed Keeper of his Majesty's Gardens, Vines, and Silkworms at Oatlands Palace, in place of his father who had died that year, and thus became head gardener to Charles I and Henrietta Maria of France. He and his father maintained a Cabinet of Curiosities known as the Ark, and added to it items collected in Virginia, including the ceremonial cloak of Chief Powhatan, an important Native American relic. Wikipedia says: “He published the contents of his father's celebrated collection as Musaeum Tradescantianum - books, coins, weapons, costumes, taxidermy, and other curiosities - dedicating the first edition to the Royal College of Physicians (with whom he was negotiating for the transfer of his botanic garden), and the second edition to the recently restored Charles II. Tradescant bequeathed his library and museum to (or some say it was swindled from him by) Elias Ashmole (1617–1692), whose name it bears as the core of the Ashmolean Museum in Oxford where the Tradescant collections remain largely intact. He was buried beside his father. The genus Tradescantia was published by Carl Linnaeus in 1753. (ref. genus Tradescantia) (Photo credit: Find-a-Grave)
  • Tra'gia: Both Gledhill in The Names of Plants and Umberto Quattrocchi in the CRC World Dictionary of Plant
      Names claim that this name was derived from the Latin name of Hieronymus Tragus (Jerome Bock) (1498-1554), a German Lutheran physician and herbalist and author of New Kreütterbuch (or Kraüterbuch, not to be confused with Leonhart Fuchs’ 1543 New Kreuterbuch), however David Hollombe has uncovered a citation from Linnaeus who authored the genus to Caspar Bauhin where in his 1623 work Pinax Theatri Botanici, page 289, he refers back to Pliny and Dioscorides in connection with the origins of the name, so this may be another example of misinformation
    having been perpetuated through the literature, but for the time being I am going with Hieronymus Bock. Many of the details of his life are unclear, for instance his birthplace and where he received his early schooling, but he was clearly interested in botany from an early age and by 1523 was head of a botanical garden owned by the Count Palatine Ludwig in Zweibrücken. He apparently attended the University of Heidelberg at least for a time. It is uncertain whether he studied medicine, philosophy, theology, or something else, but there does not seem to be any record of his having received a degree. He presumably did have some theological training. Following his attendance there he became a schoolteacher in Zweibrücken for nine years. In 1533 he received a life-time position as a Lutheran minister at the Benedictine church of St. Fabian in nearby Hornbach where he mostly stayed up to his death. He left Hornbach for a brief period in 1550 and acted as personal physician to the Landgraf Philipp II of Nassau, whose garden he is said to have supervised and to whom his Kreutterbuch was dedicated. In 1551 he returned to Hornbach, where he died three years later, probably of consumption. Wikipedia says: “The first edition of his Kreutterbuch (literally ‘plant book’) appeared in 1539 unillustrated; his stated objectives were to describe German plants, including their names, characteristics, and medical uses. Instead of following Dioscorides as was traditional, he developed his own system to classify 700 plants. Bock apparently traveled widely through the German region observing the plants for himself, since he includes ecological and distributional observations. A subsequent edition of the Kreutterbuch published in 1546was illustrated by the artist David Kandel. The grass genus Tragus (published by Albrecht von Heller in 1768) and the spurge genus Tragia (published by Linnaeus in 1753) are both said to be named after him. (ref. genus Tragia)
  • Tragopo'gon: derived from two Greek words tragos meaning "goat" and pogon meaning "beard," suggested by its prominent, feathery hairs when in seed. The genus Tragopogon was published by Carl Linnaeus in 1753. (ref. genus Tragopogon)
  • tra'gus: possibly from the Greek tragos, a part of the ear, literally "goat," or from Hieronymous Tragus, the Greek name for Jerome Bock (1498-1554), physician, scholar, and one of the three fathers of German botany. (ref. Salsola tragus)
  • trans-: beyond, through, across.
  • Transberin'gia: this genus contains a single species with two subspecies, one in northwestern North America and one in northeastern Asia, separated by the Bering Strait, hence 'trans' 'Bering.' The genus Transberingia was published by Ihsan Ali Al-Shehbaz and Steve Lawrence O'Kane in 2003. (ref. genus Transberingia)
  • transitor'ius: I am assuming that this means what the modern English word 'transitory' means, that is "of brief duration, not persistent, temporary," but it may alternatively mean "transitional or intermediate." (ref. Rumex salicifolius var. transitorius)
  • transmonta'na: beyond the mountains. (ref. Gilia transmontana)
  • transvalen'sis: of or from the Transvaal Province of South Africa. (ref. Cynodon transvalensis)
  • transver'sa: transverse, set crosswise, made at right angles. (ref. Gilia leptantha ssp. transversa, Piperia transversa)
  • trask'ae/trask'iae: after Luella Blanche (Engle) Trask (1865-1916), resident of Catalina Island from 1895 to 1915 and botanical explorer and collector who lost many specimens in the 1906 San Francisco earthquake and fire and a later fire in Avalon on Catalina Island in 1916. Born in Waterloo, Iowa, she moved with her family to Minnesota while still a youngster. Her father was a nurseryman. She seemed to have had a love of flowers from the earliest age, and recalled the story that her mother would find her in the garden at the age of 2 or 3 kissing the pansies and verbenas She studied and collected plants in the southern part of Minnesota, before eventually marrying, having a daughter, and moving to Santa Monica, California. In addition to botany, her interests included archeology, history, zoology and geology, and beyond the Channel Islands she also worked and explored in the deserts of the Southwest, the San Jacinto Mountains, Death Valley, Arizona, Utah, Colorado and Yellowstone. She carried on an extensive correspondence with Alice Eastwood and Willis Jepson. Her winter home was in Avalon, but she also had a summer refuge in Fisherman's Cove, and frequently walked the roundtrip route over the ridge trail in a day. She was described by Charles Frederick Milspaugh, former Curator of the Field Museum of Natural History in Chicago, as an "indefatigable pedestrian." (ref. Astragalus traskiae, Cercocarpus traskiae, Dudleya traskiae, Cryptantha traskiae, Eriodictyon traskiae, Mimulus traskiae)
  • Trautvetter'ia: after Ernst Rudolf von Trautvetter (1809-1889), a Baltic-German or Russian botanist who
      specialized in the flora of the Caucasus and central Asia and was author of De Echinope genere capita (1833) and the Decas plantarum novarum published in St. Petersburg, 1882. He was born in Jelgava, Latvia, the son of a German philosophy professor, teacher and author of De novo systemate botanico brevem notitiam  Ernst Christian Johann von Trautvetter. Wikipedia says: “He studied medicine and natural sciences at the University of Dorpat [in present-day Estonia.] From 1829 to 1831, he conducted botanical field trips throughout Livonia, returning to Jelgava in
    1831 as a private instructor. In 1833 he began work as an assistant at the botanical garden in Dorpat, two years later, performing similar duties at the botanical garden in St. Petersburg. In 1838 he relocated to Kiev as a professor of botany and director of the botanical garden. During his many years in Kiev, he served as university rector from 1847 to 1859. Later in his career, he returned to the botanical garden in St. Petersburg as an administrator and director. Here, he was tasked with publishing an account of the garden's history.” His first publication was entitled Ueber die Nebenblätter (1831) and was a work on plant stipules. In 1835 he received a doctor of philosophy degree from the University of Königsberg. After a brief time as director of the Agricultural Institute at Gory-Gorki (Mogilev) he returned to the Botanic Garden of St. Petersburg to become the director there. He retired in 1875, only to return to directorship in 1889, the same year in which he died. He was a prolific author, publishing among other things Skizze der Classen und Ordnungen des natürlichen Ppflanzensysterns which was hisassessment of the natural orders and classes of plant systems. He also added to the regional floras of northern Soberia and the Caspian-Caucusus region. The genus Trautvetteria was published by Friedrich Ernst Ludwig von Fischer and Carl Anton Meyer in 1835. (ref. genus Trautvetteria)
  • treleas'ei: after William Trelease (1857-1945), who was "born in Mount Vernon, New York in 1857. In his teens
      he was briefly apprenticed in a machine shop, but in 1877 he decided to enter Cornell College and study the natural sciences. After Cornell, he taught at the University of Wisconsin, and planned to study bacteriology there; but when he was offered the director's job at the St. Louis Botanical Garden, he accepted. It was an ideal position for Trelease, a gifted botanist with a genius for classifying plants. He directed the 75 acre garden for 23 years, during which time he identified and named 2500 species and varieties of flora. Trelease's botanical interests were broad: he published a
    paper on the giant cactus of Mexico in the same year that he published his findings about coastal species in Alaska. He studied apple scab, leaf blight, nematodes. [In 1899 he joined John Muir on the Harriman Alaska expedition aboard the ship Elder.] While on the Elder, he worked with the others involved in botany, collecting specimens, but tending to play second fiddle to Muir, Gilbert and the other, more talkative, scientists. He was first and foremost a scientist, not a story-teller. After the expedition, he returned to St. Louis, and eventually taught at the University of Illinois. His work includes hundreds of scholarly papers, but Trelease was not solely an academic. He published a small, inexpensive set of guides for the everyday gardener, that remain, to this day, a value for anyone who wishes to graft an apple tree, or identify and avoid poison ivy in the winter. Trelease died in 1945." He was the author of Winter Botany: An Identification Guide to Native Trees & Shrubs and his name is on Mt. Trelease in Colorado. (ref. Draba paysonii var. treleasei, Opuntia basilaris var. treleasei) (Photo credit: Herbarium of the L.H. Bailey Hortorium)
  • trem'ula: trembling, referring to foliage.
  • tremulo'ides: like the quivering poplar or quaking aspen. (ref. Populus tremuloides)
  • tri-: in compound words signifying three.
  • triacan'thos: three-spined. (ref. Gleditsea triacanthos)
  • Triad'ica: from the Greek for three, for lobes of calyx, ovary, fruit. (ref. genus Triadica)
  • triangular'is: with three angles. (ref. Pentagramma triangularis, Senecio triangularis)
  • triangulival'vis: presumably with one or more three-angled or triangular valves. (ref. Rumex triangulivalvis)
  • Trian'tha: probably the same as Trianthema, that is, three-flowered. The genus Triantha was published by John Gilbert Baker in 1879. (ref. genus Triantha)
  • Trianth'ema: from the Greek treis, "three," and anthemon, "flower." The genus Trianthema was published by John Gilbert Baker in 1879. (ref. genus Trianthema)
  • Tribo'lium: from the Greek name tribolos applied to various prickly plants. (ref. genus Tribolium)
  • tribractea'tum: three-bracted. (ref. Allium tribracteatum, Lythrum tribracteatum)
  • Trib'ulus: from the Greek tribeles or tribolos and Latin tribulus for "three-pointed, a caltrop," the shape of which is suggested by the three-pronged fruit, and referring to the caltrop, an ancient military weapon which consisted of an iron ball with projecting spikes that could be strewn on the battlefield to impede cavalry or foot soldiers. At least one spike projected upwards while others anchored the caltrop in the ground. The genus Tribulus was published by Carl Linnaeus in 1753. (ref. genus Tribulus)
  • Tricar'dia: from the Greek tri for "three" and cardia for "heart," thus "three-hearted," apparently from the structure of the calyx. The genus Tricardia was published by John Torrey in 1871. (ref. genus Tricardia)
  • tricarinat'us: three-keeled, or by extension, three-ribbed or with three angled sides. (ref. Astragalus tricarinatus)
  • tri'ceps: three-headed. (ref. Gilia cana ssp. triceps)
  • trichan'tha: three-flowered. (ref. Zeltnedra trichantha)
  • tricho-: in compound words signifying hairy or hair-like.
  • trichoca'lyx: with a hairy calyx. (ref. Eriodictyon trichocalyx var. lanatum, Eriodictyon trichocalyx var. trichocalyx, Romneya trichocalyx)
  • trichan'thum: hairy-flowered. (ref. Centaurium trichanthum)
  • trichocar'pa: hairy-fruited. (ref. Populus trichocarpa)
  • Trichocoron'is: from the Greek trichos, "hair," and koronis, "crown" referring to the pappus. The genus Trichocoronis was published by Asa Gray in 1849. (ref. genus Trichocoronis)
  • trichoma'nes: the name of a fern mentioned by Theophrastus and Dioscorides, and used by Pliny for a fern resembling Adiantum.
  • tricho'pes: hairy-footed or -stalked. (ref. Eriogonum trichopes)
  • Trichophor'um: from the Greek thrix or trichos, "hair," and phoros, "bearing, carrying," referring to the margins of the clinandrum, which is the portion of an orchid column which conceals the anther. The genus Trichophorum was published by Christiaan Hendrik Persoon in 1805. (ref. genus Trichophorum)
  • trichopo'dus: having a hairy pod. (ref. Astragalus trichopodus var. lonchus, Astragalus trichopodus var. phoxus)
  • Trichoptil'ium: from the Greek trichos, "hair," and ptilon, "feather," in allusion to the dissected pappus-paleae, i.e. the chafflike scales on the receptacle. The genus Trichoptilium was published by Asa Gray in 1859. (ref. genus Trichoptilium)
  • Trichos'tema: from trichos, "hair," and stema, "stamens," and alluding to the hair-like stamens. The genus Trichostema was published by Johan Frederik Gronovius in 1753. (ref. genus Trichostema)
  • tri'color: of three colors. (ref. Gilia tricolor, Verbena tricolor)
  • tricorna'tum: three-horned. (ref. Galium tricornatum)
  • tricuspida'ta: three-pointed.
  • tricus'pis: probably meaning the same as the above. (ref. Mentzelia tricuspis)
  • tridactylo'sa: from tri-, "three," and daktylos, "a finger, toe," thus meaning three-fingered or three-toed. (ref. Calystegia collina ssp. tridactylosa)
  • Tri'dens: three-toothed. The genus Tridens was published by Johann Jakob Roemer and Josef August Schultes in 1817. (ref. genus Tridens)
  • tridenta'ta/tridenta'tum: three-toothed. (ref. Artemisia tridentata ssp. tridentata, Artemisia tridentata ssp. vaseyana, Larrea tridentata, Mentzelia tridentata, Purshia tridentata var. glandulosa, Trifolium tridentatum)
  • Trienta'lis: one-third of a foot in height, about the height of these small plants. The genus Trientalis was published by Carl Linnaeus in 1753. (ref. genus Trientalis)
  • tri'fida/tri'fidum: cleft into three parts. (ref. Ambrosia trifida, Bouteloua trifida, Corallorhiza trifida, Galium trifidum var. subbiflorum)
  • triflor'um: three-flowered. (ref. Galium triflorum, Solanum triflorum)
  • trifolia'ta: three-leaved. (ref. Menyanthes trifoliata)
  • Trifo'lium: from the Latin meaning "three-leaved." The genus Trifolium was published by Carl Linnaeus in 1753. (ref. genus Trifolium)
  • trifor'mis: Engelmann described the species as having three varieties: var stylosus ('planta major...'), var. brachystylus ('planta minor...') and var. uniflorus ('planta minima...'). (ref. Juncus triformis)
  • trifur'ca: thrice forked. (ref. Ephedra trifurca)
  • triglochidia'tus: with three barbed bristles. (ref. Echinocereus triglochidiatus)
  • Triglo'chin: from the Greek tri, "three," and glochis, "a point," referring to the fruit of some species. The genus Triglochin was published by Carl Linnaeus in 1753. (ref. genus Triglochin)
  • Trigonel'la: a diminutive of the Latin trigonum, "triangle or three-cornered," referring to the corolla of one of the species. The genus Trigonella was published by Carl Linnaeus in 1753. (ref. genus Trigonella)
  • trigonophyl'la: possibly from trigonus, "three-cornered," and phyllon, "leaf." (ref. Nicotiana trigonophylla)
  • trigy'na/trigyn'um: with three pistils. (ref. Linum trigynum)
  • trilliifo'lia: with leaves like genus Trillium. (ref. Oxalis trilliifolia)
  • trilo'ba/trilo'bata: indicates leaves are three-lobed. (ref. Ipomoea triloba, Oxytheca trilobata)
  • trimes'tris: blooming for three months. (ref. Lavatera trimestris)
  • Trimor'pha: from the Greek treis or tria, "three," and morphe, "a shape," in reference to the flower types. The genus Trimorpha was published by Alexandre Henri Gabriel de Cassini in 1817. (ref. genus Trimorpha)
  • triner'via: with three nerves or veins. (ref. Flaveria trinervia)
  • trin'ii: after Karl Bernhard von Trinius (1778-1844), German court physician, poet and world-famous agrostologist
      who was born in Eisleben northeast of Leipzig. He founded the Botanical Museum in St. Petersburg and was personal physician and teacher of Tsar Alexander II. He was the son of a Protestant pastor. His uncle was the founder of homeopathy, Samuel Hahnemann. His father died early and his mother married a teacher named Müller from Eisleben. He studied medicine from 1792 to 1802, first in Jena, then in Halle, and finally in Leipzig. Along the way he as was the case with so many other medical people developed an interest in botany. He received a doctorate in
    1802 in Göttingen. After finishing his studies he moved to the Baltic provinces and located in several different communities. He married in 1804 and in 1808 became the personal physician of the Duchess Antoinette of Württemberg. He travelled extensively throughout Germany and Russia and from 1811 to 1815 was with the Duchess in St. Petersburg. During the period 1816 to 1822 he lived in Belarus before returning to St. Petersburg and becoming a full member of the Imperial Academy of Sciences as a result of his botanical work. In 1821 he was elected a member of the German Academy of Sciences Leopoldina. In 1824 he was appointed as the imperial personal physician and taught natural sciences to Tsar Alexander II from 1829 to 1833. At this time he was continuing to communicate with his uncle and delve into the study of homeopathy. He visited botanical collections abroad on behalf of the imperial academy in 1836 but in 1837 and 1838 he suffered several strokes and died in 1844 in St. Petersburg. He was also a poet and a prolific author, and compiled a herbarium of 4000-5000 specimens which he bequeathed to the Botanical Museum of St. Petersburg. (ref. Bromus trinii)
  • triniten'sis: of or from the area of the Trinity Mountains of northern California. (ref. Arctostaphylos trinitensis)
  • tri'odon: from the Greek for "three" and "tooth," referring to the three lobes of the petals. (ref. Aliciella triodon)
  • Triodan'is: from Greek treis, "three," and odons, "tooth," hence "three-toothed." The genus Triodanis was published by Constantine Samuel Rafinesque in 1836. (ref. genus Triodanis)
  • trio'num: Jaeger's Source-book of Biological Names and Terms gives two alternate etymologies for this name: (1) from New Latin trionum derived from the Greek trionon, the name of a malvaceous plant; and (2) from Latin Triones, referring to Ursa Major or the Big Dipper. "Ursa Major in English tradition is the Plough derived possibly from the Triones or Teriones, the Plough Oxen or Threshing Oxen of Roman fable; Cicero mentions them as the Septentriones, which later became a term for the north wind, the northern heavens and polar things in general." (From a website on Ursa Major) Since this plant is malvaceous and not particular polar or northern, the former etymology is probably correct. (ref. Hibiscus trionum)
  • tripar'tita: in three parts, having three parts. (ref. Bidens tripartita)
  • triphyl'la: three-leaved. (ref. Achlys triphylla, Lewisia triphylla)
  • triphyl'los: probably same as previous entry. (ref. Veronica triphyllos)
  • Triphysar'ia: from the Greek meaning "having three bladders" because of the lower lip pouches. The genus Triphysaria was published by Friedrich Ernst Ludwig von Fischer and Carl Anton von Meyer in 1836. (ref. genus Triphysaria)
  • Tripleurosper'mum: from the Greek treis, "three," pleura or pleuron, "rib," and sperma, "seed," referring to the achenes. The genus Tripleurospermum was published by Carl Heinrich Schultz in 1844. The genus Tripleurospermum was published by Carl Heinrich Schultz in 1844. (ref. Tripleurospermum)
  • tripo'dum: three-footed. (ref. Eriogonum tripodum)
  • Tripteroca'lyx: from the Greek tripteros, "three-winged," and kalyx, "calyx." The genus Tripterocalyx was published by William Jackson Hooker in 1853. (ref. genus Tripterocalyx)
  • triquet'ra/triquet'rum: from the Latin triquetrus meaning "having three corners, triangular." (ref. Boerhavia triquetra, Carex triquetra, Allium triquetrum)
  • trisep'alus: with three sepals. (ref. Ranunculus bonariensis var. trisepalus)
  • Triset'um: from tri, "three," and seta, "bristle, hair," referring to the three-bristled appearance of the lemma of the type species T. flavescens, which results from the presence of a bristle and two teeth. The genus Trisetum was published by Christiaan Hendrik Persoon in 1805. (ref. genus Trisetum)
  • trista'chya: three-spiked. (ref. Eleusine tristachya)
  • tris'te: dull, sad. (ref. Hieracium triste)
  • tris'tulis: diminutive of tristis, "sad." (ref. Fritillaria affinis var. tristulis)
  • trisul'ca: with three furrows. (ref. Lemna trisulca)
  • Tritel'eia: from the Greek tri, "three," and teleios, "perfect," the floral parts being in 3's. The genus Triteleia was published by David Douglas in 1830. (ref. genus Triteleia)
  • triterna'tum: triply ternate. (ref. Lomatium triternatum)
  • tritico'ides: like genus Triticum, or wheat. (ref. Leymus triticoides)
  • Trit'icum: the classical Latin name for wheat. The genus Triticum was published by Carl Linnaeus in 1753. (ref. genus Triticum)
  • triuncia'lis: the Latin uncia means one-twelfth, so this might mean three-twelfths, of unknown application. (ref. Aegilops triuncialis)
  • trivia'le/trivia'lis: common, ordinary. (ref. Alisma triviale, Poa trivialis)
  • trixa'go: Brown's Composition of Scientific Words says that this name is Latin for germander, but then -ago is a Latin substantival suffix used to indicate a resemblance or property, and trix could derive from trix (thrix), "a hair," or trixos "three-fold," inasmuch as B. trixago is a glandular-hairy plant, and it was apparently an old name for some plant possibly of the mint family. (ref. Bellardia trixago)
  • Trix'is: from the Greek trixos, "three-fold," referring to the three-cleft outer corolla lip. The genus Trixis was published by Patrick Browne in 1756. (ref. genus Trixis)
  • trochlear'is: from the Greek trochlea, "a pulley," plus -aris, "belonging or pertaining to," of uncertain application. (ref. Callitriche trochlearis)
  • trolliifo'lium: with leaves like genus Trollius in the buttercup family. (ref. Delphinium trolliifolium)
  • Tropaeo'lum: the nasturtium of gardeners, although not of botanists, and named by Linnaeus, from the Greek tropaion and the Latin tropaeum for "trophy," the manner in which the plant grew up a support reminding him of a classical trophy with round shields and golden helmets such as those hung as a sign of victory on a battlefield. The genus Tropaeolum was published by Carl Linnaeus in 1753. (ref. genus Tropaeolum)
  • Tropidocar'pum: from the Greek meaning "keel fruit," referring to the keeled capsular valves. The genus Tropidocarpum was published by William Jackson Hooker in 1836. (ref. genus Tropidocarpum)
  • troximo'ides: Jaeger's Sourcebook relates the root trox with trog, from the Greek trogo, "to gnaw" and trogos, "a gnawer, weevil", and Brown's Composition of Scientific Words says that troxalis is the Latin word for grasshopper or cricket, which are gnawing insects. Another piece of evidence for this interpretation is a web listing that I found of the species Microseris troximoides (synonym for Nothocalais troximoides) for which the common name weevil microseris is given. (ref. Nothocalais troximoides)
  • tru'ei: after Gordon Haines True, Jr. (1908-1984), son of livestock scientist and Professor of Animal Husbandry at UC Davis. True, Jr. graduated from Davis with a degree in Animal Science and received his MS in parasitology from Berkeley in 1931. He first worked as a parasitologist for the California State Department of Fish and Game and then the Bureau of Game Conservation in San Francisco until 1946. After being injured in a car accident, he moved to Nevada to work a dairy ranch and collect local plants. He moved back to California in 1963, two years later becoming a Research Associate in the Botany Department at the California Academy of Sciences. (ref. Arctostaphylos mewukka ssp. truei)
  • trunca'ta/trunca'tum/trunca'tus: abruptly cut off. (ref. Atriplex truncata, Horkelia truncata, Pellaea truncata, Lupinus truncatus)
  • truxillen'sis: apparently a reference to the city of Trujillo in Peru, since C. truxillensis was first described from Peru. Trujillo lies on the Pan-American Highway NNW of Lima, in the coastal desert region of the Moche River Valley, and is a commercial, educational, cultural and transportation center. It is the 2nd oldest Spanish city in Peru and was founded in 1534 by Diego de Almagro and elevated to city status the following year by Francisco Pizarro who named it after his birthplace in Spain. Trujillo was visited by Alexander von Humboldt during his epic excursion across Central and South America (1799-1804) in the course of which he travelled some 6000 miles on foot and horseback and by canoe. (ref. Cressa truxillensis)
  • Tsu'ga: from a Japanese name for their native hemlocks. The genus Tsuga was published by Élie Abel Carrière in 1855. ref. genus Tsuga)
  • tsugen'se: of hemlocks. (ref. Arceuthobium tsugense)
  • tu'berans/tuberas'cens: becoming swollen or tuberous.
  • Tuberar'ia: from the thickened, tuber-like swellings on the roots of the type species. The genus Tuberaria was published by Édouard Spach in 1836. (ref. genus Tuberaria)
  • tubero'sa: tuberous, alluding to the fact that the rhizomes of some species have tubers. (ref. Anemone tuberosa, Sanicula tuberosa, Scutellaria tuberosa)
  • tubiflor'a: with tubular flowers.
  • tubulo'sa: tubular, pipe-like. (ref. Swertia tubulosa)
  • Tuctor'ia: apparently an anagram of the genus name Orcuttia. The genus Tuctoria was published by John Raymond Reeder in 1982. (ref. genus Tuctoria)
  • tularen'se/tularen'sis: of or from the Tulare area of the San Joaquin Valley. (ref. Ribes tularense, Atriplex tularensis)
  • Tul'ipa: Latinized version of the Turkish tulbend, a turban. The genus Tulipa was published by Carl Linnaeus in 1753. (ref. genus Tulipa)
  • tumes'cens: swollen. (ref. Arctostaphylos parryana ssp. tumescens)
  • tumulico'la: a dweller of mounds and hillocks. (ref. Carex tumulicola)
  • tumulo'sa: from the Latin tumulus, "a mound or hillock," and meaning "swollen." (ref. Cryptantha tumulosa)
  • tuolumnen'se: from Tuolumne County. (ref. Allium tuolumnense)
  • turbina'ta: shaped like a spinning top. (ref. Abronia turbinata)
  • turbinel'la: may mean the same as turbinata (see above) although possibly in diminutive form. (ref. Quercus turbinella)
  • turionif'era: bearing turions, which are small shoots or buds. (ref. Lemna turionifera)
  • Turric'ula: Latin for "little tower." The genus Turricula was published by James Francis Macbride in 1917. (ref. genus Turricula)
  • Turrit'is: according to the Jepson Manual and Jaeger's Source-Book of Biological Names and Terms, this generic epithet is from the Latin turris for "tower" or turritus, "furnished with towers," for the orientation of the overlapping leaves and fruits, giving the plant a pyramidal shape. The genus Turritis was published by Carl Linnaeus in 1753. (ref. genus Turritis)
  • Twisselmann'ia/twisselmann'ii: after Ernest Christian Twisselmann (1917-1972), cattle rancher, authority
      on southern San Joaquin Valley flora, and author of The Flora of Kern County. He lwas born and ived nearly all his life in Cholame in San Luis Obispo County east of Paso Robles. His family had large holdings in the Cholame Valley and Temblor Range. He graduated from the University of California, Berkeley, and shortly after began cattle ranching which he continued until his death. An outbreak of nitrate poisoning in 1952 that caused extensive cattle losses in parts of California triggered an interest in Twisselman to discover the plants causing the outbreak, and this led him
    into botany. He learned the mechanics of collecting plants and how to identify them from John Thomas Howell and in 1956 published a flora of the Temblor Range region. His focus expanded to Kern County and his Kern County book was published in 1968. As early as 1965 he began field work with the cooperation of USFS on the Kern Plateau in the southern Sierra Nevada of Tulare and Kern Counties. He built up a personal herbarium of about 20,000 specimens which was eventually held by the California Academy of Sciences. Most of this information comes from an obituary in Madroño, Vol. 22, No. 8, October 1974. The genus Twisselmannia was published by Ihsan Ali Al-Shehbaz in 1999. (ref. genus Twisselmannia, also Eriogonum twisselmannii, Nemacladus twisselmannii) (Photo credit: A Flora of Kern County)
  • Ty'pha: the Greek name for this plant. The genus Typha was published by Carl Linnaeus in 1753. (ref. genus Typha)

Mt. San Gorgonio from Joshua Tree National Park
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